It is the dominant and widely ranging species which vary most frequently and vary most, and varieties are often at first local--both causes rendering the discovery of intermediate links in any one formation less likely.

Local varieties will not spread into other and distant regions until they are considerably modified and improved; and when they have spread, and are discovered in a geological formation, they appear as if suddenly created there, and will be simply classed as new species.

Most formations have been intermittent in their accumulation; and their duration has probably been shorter than the average duration of specific forms.