Possibly these several differences may be connected with the different flow of nutriment towards the central and external flowers.

We know, at least, that with irregular flowers those nearest to the axis are most subject to peloria, that is to become abnormally symmetrical.

I may add, as an instance of this fact, and as a striking case of correlation, that in many pelargoniums the two upper petals in the central flower of the truss often lose their patches of darker colour; and when this occurs, the adherent nectary is quite aborted, the central flower thus becoming peloric or regular.

When the colour is absent from only one of the two upper petals, the nectary is not quite aborted but is much shortened.

With respect to the development of the corolla, Sprengel's idea that the ray-florets serve to attract insects, whose agency is highly advantageous, or necessary for the fertilisation of these plants, is highly probable; and if so, natural selection may have come into play.